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Home > Curated Information Page > Cst Curation Set #9737
Possemato A (2010) CST Curation Set: 9737; Year: 2010; Biosample/Treatment: cell line, H1650/Phenformin; Disease: non-small cell lung cancer; SILAC: -; Specificities of Antibodies Used to Purify Peptides prior to LCMS: RXXp[ST]
This page summarizes selected information from the record referenced above and curated into PhosphoSitePlus®, a comprehensive online resource for the study of protein post-translational modifications (NAR, 2015, 43:D512-20). To learn more about the scope of PhosphoSitePlus®, click here.
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T302-p - 53BP1 (human)
Modsite: PEPEVLstQEDLFDQ SwissProt Entrez-Gene
Orthologous residues
53BP1 (human): T302‑p, 53BP1 iso2 (human): T307‑p, 53BP1 iso3 (human): T307‑p, 53BP1 (mouse): T305‑p, 53BP1 iso10 (mouse): T305‑p, 53BP1 (rat): T307‑p, 53BP1 (cow): T292‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S46-p - ALDOA (human)
Modsite: sIAkRLQsIGtENtE SwissProt Entrez-Gene
Orthologous residues
ALDOA (human): S46‑p, ALDOA (mouse): S46‑p, ALDOA (rat): S46‑p, ALDOA (rabbit): S46‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S344-p - CCNYL1 (human)
Modsite: AAMRRsFsADNFIGI SwissProt Entrez-Gene
Orthologous residues
CCNYL1 (human): S344‑p, CCNYL1 iso2 (human): S293‑p, CCNYL1 (mouse): S276‑p, CCNYL1 (rat): S275‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S78-p - CEP131 (human)
Modsite: NNLRRsNsttQVSQP SwissProt Entrez-Gene
Orthologous residues
CEP131 (human): S78‑p, CEP131 iso2 (human): S78‑p, CEP131 (mouse): S78‑p, CEP131 (rat): S78‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S404-p - CEP152 iso1 (human)
Modsite: LGIHPSDsEGELNIE SwissProt Entrez-Gene
Orthologous residues
CEP152 (human): S497‑p, CEP152 iso1 (human): S404‑p, CEP152 (mouse): S487‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S131-p - CGN (human)
Modsite: GKLLRsHsQAsLAGP SwissProt Entrez-Gene
Orthologous residues
CGN (human): S131‑p, CGN iso2 (human): S131‑p, CGN (mouse): S131‑p, CGN iso2 (mouse): S131‑p, CGN (rat): S132‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S600-p - CLASP1 (human)
Modsite: GsLQRsRsDIDVNAA SwissProt Entrez-Gene
Orthologous residues
CLASP1 (human): S600‑p, CLASP1 iso2 (human): S600‑p, CLASP1 iso3 (human): S600‑p, CLASP1 iso5 (human): S600‑p, CLASP1 (mouse): S600‑p, CLASP1 iso3 (mouse): S600‑p, CLASP1 (rat): S542‑p, CLASP1 iso5 (rat): S542‑p, CLASP1 (cow): S536‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T551-p - CTNNB1 (human)
Modsite: HQDtQRRtsMGGtQQ SwissProt Entrez-Gene
Orthologous residues
CTNNB1 (human): T551‑p, CTNNB1 (mouse): T551‑p, CTNNB1 (rat): T551‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S252-p - CTNND1 (human)
Modsite: MEGyrAPsRQDVyGP SwissProt Entrez-Gene
Orthologous residues
CTNND1 (human): S252‑p, CTNND1 iso2 (human): S252‑p, CTNND1 iso3 (human): S252‑p, CTNND1 iso4 (human): S252‑p, CTNND1 iso10 (human): S198‑p, CTNND1 iso12 (human): S198‑p, CTNND1 iso22 (human): S151‑p, CTNND1 (mouse): S252‑p, CTNND1 iso3 (mouse): S252‑p, CTNND1 (rat): S252‑p, CTNND1 (cow): S252‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S268-p - CTNND1 (human)
Modsite: PQVRVGGssVDLHRF SwissProt Entrez-Gene
Orthologous residues
CTNND1 (human): S268‑p, CTNND1 iso2 (human): S268‑p, CTNND1 iso3 (human): S268‑p, CTNND1 iso4 (human): S268‑p, CTNND1 iso10 (human): S214‑p, CTNND1 iso12 (human): S214‑p, CTNND1 iso22 (human): S167‑p, CTNND1 (mouse): S268‑p, CTNND1 iso3 (mouse): S268‑p, CTNND1 (rat): S268‑p, CTNND1 (cow): S268‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S423-p - DNAJA5 (human)
Modsite: DTNLNQDsAKELEDs SwissProt Entrez-Gene
Orthologous residues
DNAJA5 (human): S423‑p, DNAJA5 (mouse): N423‑p, DNAJA5 (rat): N186‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S678-p - EPB41L1 (human)
Modsite: EsGGIEDsPDRGACs SwissProt Entrez-Gene
Orthologous residues
EPB41L1 (human): S678‑p, EPB41L1 iso2 (human): S604‑p, EPB41L1 iso4 (human): , EPB41L1 (mouse): S677‑p, EPB41L1 iso4 (mouse): , EPB41L1 iso5 (mouse): S677‑p, EPB41L1 (rat): S677‑p, EPB41L1 iso2 (rat): S1349‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S685-p - EPB41L1 (human)
Modsite: sPDRGACstPDMPQF SwissProt Entrez-Gene
Orthologous residues
EPB41L1 (human): S685‑p, EPB41L1 iso2 (human): S611‑p, EPB41L1 iso4 (human): , EPB41L1 (mouse): S684‑p, EPB41L1 iso4 (mouse): , EPB41L1 iso5 (mouse): S684‑p, EPB41L1 (rat): S684‑p, EPB41L1 iso2 (rat): S1356‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T686-p - EPB41L1 (human)
Modsite: PDRGACstPDMPQFE SwissProt Entrez-Gene
Orthologous residues
EPB41L1 (human): T686‑p, EPB41L1 iso2 (human): T612‑p, EPB41L1 iso4 (human): , EPB41L1 (mouse): T685‑p, EPB41L1 iso4 (mouse): , EPB41L1 iso5 (mouse): T685‑p, EPB41L1 (rat): T685‑p, EPB41L1 iso2 (rat): T1357‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S295-p - FNBP1L (human)
Modsite: QHIyRtIsDGtIsAS SwissProt Entrez-Gene
Orthologous residues
FNBP1L (human): S295‑p, FNBP1L iso2 (human): S295‑p, FNBP1L iso4 (human): S295‑p, FNBP1L (mouse): S295‑p, FNBP1L (rat): S295‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S87-p - HN1 (human)
Modsite: sGLQRRNssEAssGD SwissProt Entrez-Gene
Orthologous residues
HN1 (human): S87‑p, HN1 iso2 (human): S87‑p, HN1 (mouse): S87‑p, HN1 (rat): S82‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S88-p - HN1 (human)
Modsite: GLQRRNssEAssGDF SwissProt Entrez-Gene
Orthologous residues
HN1 (human): S88‑p, HN1 iso2 (human): S88‑p, HN1 (mouse): S88‑p, HN1 (rat): S83‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S1021-p - ITGA4 (human)
Modsite: QEENRRDsWsyINSk SwissProt Entrez-Gene
Orthologous residues
ITGA4 (human): S1021‑p, ITGA4 (mouse): S1028‑p, ITGA4 (rat): S1025‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S10-p - K19 (human)
Modsite: sySYrQssAtssFGG SwissProt Entrez-Gene
Orthologous residues
K19 (human): S10‑p, K19 (mouse): S10‑p, K19 (rat): S10‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S466-p - KLC3 (human)
Modsite: AGMkRAMsLNtLNVD SwissProt Entrez-Gene
Orthologous residues
KLC3 (human): S466‑p, KLC3 iso2 (human): S465‑p, KLC3 (mouse): S467‑p, KLC3 (rat): S467‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T308-p - KNOP1 (human)
Modsite: GDPWKEEtDtDLEVV SwissProt Entrez-Gene
Orthologous residues
KNOP1 (human): T308‑p, KNOP1 (mouse): S312‑p, TSG118 iso2 (mouse): S152‑p, KNOP1 (rat):
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S243-p - LPIN2 (human)
Modsite: PQTACPKsDsELEVK SwissProt Entrez-Gene
Orthologous residues
LPIN2 (human): S243‑p, LPIN2 (mouse): S243‑p, LPIN2 (rat): S243‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S224-p - LPIN3 (human)
Modsite: GELTSPKsDSELEVR SwissProt Entrez-Gene
Orthologous residues
LPIN3 (human): S224‑p, LPIN3 iso2 (human): S225‑p, LPIN3 (mouse): S218‑p, LPIN3 (rat): S217‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S394-p - MISP (human)
Modsite: PGLRRALssDsILsP SwissProt Entrez-Gene
Orthologous residues
MISP (human): S394‑p, MISP (mouse): S364‑p, MISP (rat): S366‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S18-p - PACT (human)
Modsite: PPLEREDsGtFsLGK SwissProt Entrez-Gene
Orthologous residues
PACT (human): S18‑p, PACT (mouse): S18‑p, PACT (rat): S18‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S578-p - PIMT (human)
Modsite: EKCQSVSsAGELETE SwissProt Entrez-Gene
Orthologous residues
PIMT (human): S578‑p, PIMT (mouse): S572‑p, PIMT (rat): S571‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S591-p - PKCI (human)
Modsite: yINPLLMsAEECV__ SwissProt Entrez-Gene
Orthologous residues
PKCI (human): S591‑p, PKCI (mouse): S590‑p, PKCI (rat): S591‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S753-p - PRICKLE2 (human)
Modsite: GQCPRtVsDLALQNA SwissProt Entrez-Gene
Orthologous residues
PRICKLE2 (human): S753‑p, PRICKLE2 (mouse): S810‑p, PRICKLE2 (rat): S756‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S407-p - RABEP1 (human)
Modsite: DGLRRAQstDsLGts SwissProt Entrez-Gene
Orthologous residues
RABEP1 (human): S407‑p, RABEP1 iso2 (human): S407‑p, RABEP1 (mouse): S407‑p, RABEP1 (rat): S407‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T408-p - RABEP1 (human)
Modsite: GLRRAQstDsLGtsG SwissProt Entrez-Gene
Orthologous residues
RABEP1 (human): T408‑p, RABEP1 iso2 (human): T408‑p, RABEP1 (mouse): T408‑p, RABEP1 (rat): T408‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S410-p - RABEP1 (human)
Modsite: RRAQstDsLGtsGsL SwissProt Entrez-Gene
Orthologous residues
RABEP1 (human): S410‑p, RABEP1 iso2 (human): S410‑p, RABEP1 (mouse): S410‑p, RABEP1 (rat): S410‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S295-p - RALY (human)
Modsite: sEEELEHsQDtDADD SwissProt Entrez-Gene
Orthologous residues
RALY (human): S295‑p, RALY iso2 (human): S279‑p, RALY iso3 (human): S296‑p, RALY (mouse): S301‑p, RALY iso2 (mouse): S285‑p, RALY (rat): S284‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T298-p - RALY (human)
Modsite: ELEHsQDtDADDGAL SwissProt Entrez-Gene
Orthologous residues
RALY (human): T298‑p, RALY iso2 (human): T282‑p, RALY iso3 (human): T299‑p, RALY (mouse): T304‑p, RALY iso2 (mouse): T288‑p, RALY (rat): T287‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S721-p - SASH1 (human)
Modsite: sQGLsGCsPRDsGCy SwissProt Entrez-Gene
Orthologous residues
SASH1 (human): S721‑p, SASH1 (mouse): S714‑p, SASH1 (rat): S711‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S113-p - TEBP (human)
Modsite: WKDWEDDsDEDMsNF SwissProt Entrez-Gene
Orthologous residues
TEBP (human): S113‑p, TEBP iso3 (human): S113‑p, TEBP (mouse): S113‑p, TEBP iso4 (mouse): , TEBP (rat): S113‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S344-p - TMF1 (human)
Modsite: RsVsEINsDDELsGK SwissProt Entrez-Gene
Orthologous residues
TMF1 (human): S344‑p, TMF1 (mouse): S340‑p, TMF1 (rat): S341‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S186-p - TOMM34 (human)
Modsite: ATKNRVPsAGDVEkA SwissProt Entrez-Gene
Orthologous residues
TOMM34 (human): S186‑p, TOMM34 (mouse): S186‑p, TOMM34 iso2 (mouse): S186‑p, TOMM34 (rat): S186‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S296-p - TRIP10 (human)
Modsite: QPMNRAPsDssLGtP SwissProt Entrez-Gene
Orthologous residues
TRIP10 (human): S296‑p, TRIP10 iso2 (human): S296‑p, TRIP10 iso3 (human): S296‑p, TRIP10 (mouse): S296‑p, TRIP10 (rat): S296‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S299-p - TRIP10 (human)
Modsite: NRAPsDssLGtPsDG SwissProt Entrez-Gene
Orthologous residues
TRIP10 (human): S299‑p, TRIP10 iso2 (human): S299‑p, TRIP10 iso3 (human): S299‑p, TRIP10 (mouse): S299‑p, TRIP10 (rat): S299‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

T302-p - TRIP10 (human)
Modsite: PsDssLGtPsDGRPE SwissProt Entrez-Gene
Orthologous residues
TRIP10 (human): T302‑p, TRIP10 iso2 (human): T302‑p, TRIP10 iso3 (human): T302‑p, TRIP10 (mouse): T302‑p, TRIP10 (rat): T302‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human

S19-p - ZNRF2 (human)
Modsite: NGRTRAysGsDLPSS SwissProt Entrez-Gene
Orthologous residues
ZNRF2 (human): S19‑p, ZNRF2 (mouse): S18‑p, ZNRF2 (rat): S18‑p, ZNRF2 (cow): S19‑p
Characterization
Methods used to characterize site in vivo mass spectrometry
Disease tissue studied:  lung cancer, non-small cell lung cancer
Relevant cell lines - cell types - tissues:  NCI-H1650 (pulmonary)
Cellular systems studied:  cell lines
Species studied:  human